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Originally Posted by rowan
I watched an entire courting ritual over a neighbour's fence when I was younger. About 5 male cats all vying for the "affections" of the sole female... the dominant one got the prize in the end.
I read somewhere that a cat penis has some sort of barb that helps it stay inside, which is why the female cats shriek... 
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I think the barb scratches the inside of the female cat's vagina, causing the pain (and the howl) and triggering ovulation.
But I could be wrong. It's been a long time since I took biology lessons.