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It's a different business model, and I have never seen the inside or a recent tour for an Adult.com site (and the examples below has nothing to do with their program whatsoever), but there are many ways programs monetize traffic, both their own and affiliate generated ...
exits
popups
cross sales
upsales
traffic leaks
malware
selling of information
It is possible (and beneficial) for any program with high member retention to pay persign up. It would be in the programs best interest to offer it. Some of the above mentioned items (like inter program upsales, cross sales and popups) can possibly be credited to affilaites as well anyways.
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